2022 in paleobotany

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This article records new taxa of fossil plants that are scheduled to be described during the year 2022, as well as other significant discoveries and events related to paleobotany that are scheduled to occur in the year 2022.

Lycopodiopsida

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Annalepis ordosensis[1]	Sp. nov	In press	Deng in Deng et al.	Middle Triassic (Ladinian)	Ordos Basin	China	A lycopsid.
Lycopodicaulis[2]	Gen. et sp. nov	Valid	Herrera et al.	Early Cretaceous (late Barremian–early Aptian)	Huolinhe Formation	China	A member of the family Lycopodiaceae. Genus includes new species L. oellgaardii.
Selaginella cretacea[3]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A species of Selaginella.

Ferns and fern allies

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Coniopteris antarctica[4]	Sp. nov	In press	Trevisan et al.	Late Cretaceous		Antarctica
Discosoropteris[5]	Gen. et 2 sp. nov	In press	Pšenička et al.	Carboniferous (Pennsylvanian)	Kladno Formation	Czech Republic	A leptosporangiate fern. Genus includes new species D. chlupatum and D. zlatkokvacekii.
Dryopterites beishanensis[6]	Sp. nov	In press	Ren & Sun in Ren et al.	Early Cretaceous	Chijinbao Formation	China
Phyllotheca douroensis[7]	Sp. nov		Barbosa et al.	Carboniferous (Gzhelian)	Douro Carboniferous Basin	Portugal	A member of Equisetales.

Conifers

Cheirolepidiaceae

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Frenelopsis antunesii[8]	Sp. nov	In press	Mendes & Kvaček	Early Cretaceous (Aptian–Albian)	Figueira da Foz Formation	Portugal	A member of the family Cheirolepidiaceae.

Cupressaceae

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Cupressinanthus klebsii[9]	Sp. nov	In press	Sadowski, Schmidt & Kunzmann	Eocene		Europe (Baltic Sea region)	Cupressaceous pollen cone.
Patagotaxodia[10]	Gen. et sp. nov	In press	Andruchow-Colombo et al.	Late Cretaceous (Maastrichtian)	Lefipán Formation	Argentina	A member of the family Cupressaceae. Genus includes new species P. lefipanensis.

Other conifers

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Aciphyllum[11]	Gen. et sp. nov		Barbacka & Górecki in Barbacka et al.	Early Jurassic (Hettangian)	Zagaje Formation	Poland	A needle leaf similar to the leaves of Pinus. Genus includes new species A. triangulatum.

Conifer research

A study on the pattern of conifer turnover across the Cretaceous-Paleogene boundary in the Raton and Denver basins (Colorado, United States) is published by Berry (2022).[12]

Flowering plants

Laurales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Catula[13]	Gen. et sp. nov	Valid	Maccracken et al.	Late Cretaceous (Campanian)	Kaiparowits Formation	United States ( Utah)	A member of the family Lauraceae. Genus includes new species C. gettyi.
Cryptocaryoxylon irregularis[14]	Sp. nov	Valid	Akkemik, Iamandei & Çelik	Early Miocene	Hançili Formation	Turkey	Fossil wood of a member of the family Lauraceae.

Lilioid monocots

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Pandanus estellae[15]	Sp. nov	Valid	Rozefelds et al.	Oligocene		Australia	A species of Pandanus.

Commelinid monocots

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Sabalites ghughuaensis[16]	Sp. nov	In press	Kumar, Hazra & Khan in Kumar et al.	Late Cretaceous-Paleocene (Maastrichtian-Danian)	Deccan Intertrappean Beds	India	A member of the family Arecaceae belonging to the subfamily Coryphoideae.
Sabalites umariaensis[16]	Sp. nov	In press	Kumar, Hazra & Khan in Kumar et al.	Late Cretaceous-Paleocene (Maastrichtian-Danian)	Deccan Intertrappean Beds	India	A member of the family Arecaceae belonging to the subfamily Coryphoideae.

Monocot research

Leaf fossils of costapalmate-palms belonging to the genus Sabalites are described from the ?Santonian–Campanian Belly River Group, Campanian Foremost Formation (Alberta, Canada) and Maastrichtian Frenchman Formation (Saskatchewan, Canada) by Greenwood, Conran & West (2022), who interpret the studied fossils as constraining climate reconstructions for the Late Cretaceous high mid-latitudes of North America (c. 55° N) to exclude significant freezing episodes; the authors also transfer the Late Cretaceous species "Geonomites" imperialis to the genus Phoenicites, and reassess Sabalites carolinensis as more likely to be Campanian than Coniacian–Santonian in age.[17]
A study on the impact of the absence of megaherbivores in the aftermath of the Cretaceous–Paleogene extinction event on the evolution of palms is published by Onstein, Kissling & Linder (2022).[18]

Proteales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Langeranthus[19]	Gen et sp nov	In press	Huegele & Manchester	Eocene Ypresian	Klondike Mountain Formation	USA Washington	A platanaceous flowering head. The type species is L. dillhoffiorum
Meliosma eosinica[20]	Sp. nov		Moiseeva, Kodrul & Jin in Moiseeva et al.	Late Eocene	Huangniuling Formation	China	A species of Meliosma.
Nelumbo fujianensis[21]	Sp. nov	In press	Dong et al.	Miocene	Fotan Group	China	A species of Nelumbo.
Platimeliphyllum durhamensis[22]	Comb nov	in press	(Wolfe)	Late Eocene	Puget Group "Upper Fultonian" Loc 9832	USA Washington	A platanaceous leaf. Moved from "Fothergilla" durhamensis (1968).
Platimeliphyllum fushunensis[22]	Comb nov	in press	(Chen)	Eocene	Fushun Formation	China	A platanaceous leaf. Moved from "Betula" fushunensis.

Protealean research

Redescription of the Okanagan Highlands genus Langeria with description of associated stipules and reproductive structures plus formal reassignment of the genus to Platanaceae by Huegele & Manchester is published.[19]

Ranunculales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Santaniella[23]	Gen. et 2 sp. nov		Gobo et al.	Early Cretaceous (Barremian-Aptian)	Crato Formation	Brazil	A member or a relative of the family Ranunculaceae. Genus includes new species S. lobata and S. acuta.

Aquifoliales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Ilex antiquorum[24]	Nom. nov	Valid	Doweld	Late Cretaceous (Maastrichtian)		Germany	A holly; a replacement name for Ilex antiqua Knobloch & Mai (1986).
Ilex myricina[24]	Nom. nov	Valid	Doweld	Miocene (Messinian)		Italy	A holly; a replacement name for Ilex myricoides Massalongo (1858).

Dipsacales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Sambucus heqingensis[25]	Sp. nov	In press	Huang & Zhou in Huang et al.	Late Pliocene	Heqing Basin	China	A species of Sambucus.

Fabales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Cercioxylon mediterraneum[14]	Sp. nov	Valid	Akkemik, Iamandei & Çelik	Early Miocene	Hançili Formation	Turkey	Fossil wood of a member of the family Fabaceae.
Paleobowdichia[26]	Gen. et comb. nov	Valid	Herendeen et al.	Latest Paleocene to late early Eocene	Lamar River Formation	United States ( Colorado Wyoming)	A member of Papilionoideae; a new genus for "Acacia" lamarensis Knowlton (1899).
Tobya[26]	Gen. et comb. nov	Valid	Herendeen et al.	Eocene	Cockfield Formation	United States ( Kentucky Tennessee)	A member of Papilionoideae; a new genus for "Diplotropis" claibornensis Herendeen & Dilcher (1990).

Fabalean research

New fossil material of members of the genus Bauhinia is described from the Eocene of the Puyang Basin (China) by Jia et al. (2022), who interpret their findings as the earliest reliable fossil records of Bauhinia in Asia.[27]

Fagales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Myricoxylon doganyurtensis[14]	Sp. nov	Valid	Akkemik, Iamandei & Çelik	Early Miocene		Turkey	Fossil wood of a member of the family Myricaceae.

Malpighiales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Belenocarpa[28]	Gen. et comb. nov	Valid	Hamersma et al.	Early Oligocene		Peru	A member of the family Euphorbiaceae; a new genus for "Jatropha" tertiara Berry.

Malvales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Thespesia neopopulnea[29]	Sp. nov	In press	Hazra, Mahato & Khan in Hazra et al.	Pliocene	Rajdanda Formation	India	A species of Thespesia.

Malvalean research

A study on the evolutionary history of Dipterocarpaceae, as indicated by biogeography of pollen fossils from Africa and India, molecular data and fossil amber records, is published by Bansal et al. (2022).[30]

Oxalidales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Ceratopetalum suciensis[31]	Sp nov	In press	Tang, Smith, & Atkinson	Late Cretaceous Campanian	Cedar District Formation	United States ( Washington)	A Cunoniaceous species.

Oxalidalean research

Tand, Smith, and Atkinson describe the first North American instance of the previously Paleo-Antarctic Rainforest Lineage Cunoniaceae fruits from Sucia Island. Previously considered solely a Gondwanan family, the new species indicate a complex geographic history for the group.[31]

Rosales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Eophylica[32]	Gen. et sp. nov	Valid	Shi et al.	Cretaceous	Burmese amber	Myanmar	A rhamnaceous floral morphotaxon. The type species is E. priscastellata.
Ficoxylon fusiforme[33]	Sp. nov	Valid	El-Noamani	Late Cretaceous	Taref Formation	Egypt	A member of the family Moraceae.
Ficus fujianensis[34]	Sp. nov	In press	Dong et al.	Miocene		China	A species of Ficus.
Ficus zhangpuensis[34]	Sp. nov	In press	Dong et al.	Miocene		China	A species of Ficus.
Phylica piloburmensis[32]	Sp. nov	Valid	Shi et al.	Cretaceous	Burmese amber	Myanmar	A species of Phylica.

Sapindales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Barkleya[35]	Gen. et comb. nov	In press	Manchester & Judd	Eocene		United States	A member of Anacardiaceae. Genus includes "Anacardites" schinoloxus Brown (1929).
Canarium haominiae[36]	Sp. nov	In press	Yin et al.	Miocene		China	A species of Canarium.
Koelreuteria kvacekii[37]	Sp. nov	Valid	Chen, Del Rio & Su in Chen et al.	Eocene	Niubao Formation	China	A species of Koelreuteria.

Other angiosperms

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Covidifructus[38]	Gen. et sp. nov	Valid	Heřmanová et al.	Late Cretaceous (late Turonian-Santonian)	Klikov Formation	Czech Republic	An angiosperm fruit of uncertain affinities, with similarities to the family Dilleniaceae. The type species is C. multicarpellatus.
Florigerminis[39]	Gen. et sp. nov	In press	Cui et al.	Middle-Late Jurassic	Jiulongshan Formation	China	A possible flower bud. The type species is F. jurassica. First announced online 2021, Final article published in 2022.	Florigerminis jurassica

General angiosperm research

A preliminary report on a new fossil angiosperm flora of the Lesvos Petrified Forest at Akrocheiras east of Sigri on Lesbos, Greece is given by Kafetzidou et al. Preliminary taxa identifications are given and commentary on the climactic implications are made.[40]
A study aiming to determine the relationship between past atmospheric CO₂ and temperature fluctuations and the shifts in diversification rates of Poaceae and Asteraceae is published by Palazzesi et al. (2022).[41]

Other plants

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Europoxylon garapensis[42]	Sp. nov	In press	Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A gymnosperm.
Kimuriella[43]	Gen. et sp. nov	Valid	Pott & Takimoto	Late Jurassic (Oxfordian)	Tochikubo Formation	Japan	A member of Bennettitales. Genus includes new species K. densifolia.
Komlopteris distinctiva[11]	Sp. nov		Barbacka in Barbacka et al.	Early Jurassic (Hettangian)	Zagaje Formation	Poland	Cuticle of a seed fern.
Lesleya ceriacoi[44]	Sp. nov	In press	Correia et al.	Carboniferous (Gzhelian)	Douro Carboniferous Basin	Portugal	An early gymnosperm.
Paragigantopteris[45]	Gen. et sp. nov	In press	Ma et al.	Permian (Wuchiapingian)	Lungtan Formation	China	A gigantopterid. Genus includes new species P. qingloongensis.
Pauthecophyton hezhangensis[46]	Sp. nov		Wang et al.	Devonian (Pragian-Emsian)	Danlin Formation	China	A euphyllophyte of uncertain affinities.
Polycanaloxylon[42]	Gen. et sp. nov	In press	Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A gymnosperm. Genus includes new species P. merlottii.
Taungurungia[47]	Gen. et sp. nov	Valid	McSweeney, Shimeta & Buckeridge	Devonian (Pragian–Emsian)	Norton Gully Sandstone Formation	Australia	A plant of uncertain affinities, similar to members of Zosterophyllopsida. Genus includes new species T. garrattii.
Teyoua[48]	Gen. et sp. nov	In press	Huang, Liu & Xue	Devonian (probably Pragian)	Mangshan Group	China	A polysporangiate land plant. Genus includes new species T. antrorsa.
Vetiplanaxis obtusus[49]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A moss belonging to the group Hypnodendrales.
Xadzigacalix[50]	Gen. et sp. nov	In press	Klymiuk, Rothwell & Stockey	Early Cretaceous		Canada ( British Columbia)	A gymnosperm of uncertain phylogenetic placement, possibly having affinities with gnetophytes or angiosperms. Genus includes new species X. quatsinoensis.

Other plant research

A study on the Paleocene charophyte flora from the South Gobi area in the Junggar Basin (China) and on the Paleogene fossil record of charophytes is published by Cao et al. (2022), who interpret their findings as evidence of the dispersal of charophyte lineages from Asia to Europe in the middle to late Eocene, possibly facilitated by waterbirds.[51]
New specimens of Radula heinrichsii, providing new information on the morphology of this liverwort, are described from the Cretaceous Burmese amber by Wang et al. (2022).[52]
The first comprehensive crown reconstruction of Medullosa stellata var. typica, based on data from a specimen from the Chemnitz petrified forest (Germany), is presented by Luthardt et al. (2022).[53]
Fossil material of Rhabdotaenia is reported from the Permian Umm Irna Formation (Jordan) by Blomenkemper et al. (2022), representing the northernmost occurrence of this Gondwanan leaf type reported to date.[54]

Palynology

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Volkheimerites[55]	Gen. et sp. nov	In press	Narváez et al.	Paleocene (Danian)	Salamanca Formation	Argentina	Pollen of a flowering plant. The type species is V. labyrinthus.
Yezopollis[56]	Gen. et sp. nov	In press	Legrand, Yamada & Nishida	Late Cretaceous (Cenomanian–Turonian)	Mikasa Formation	Japan	A Normapolles-type flowering plant pollen. Genus includes new species Y. mikasaensis.

Research

A study on the biodiversity of land plants at the equator during their first major diversification in the Late Silurian–Early Devonian is published by Wellman et al. (2022).[57]
A study on the evolution of heterospory during the Devonian is published by Leslie & Bonacorsi (2022).[58]
New Oligocene flora is described from the Dong Ho Formation (Vietnam) by Huang et al. (2022), who interpret the studied fossils as evidence of long-term environmental, floristic and vegetational stability in this region since the Paleogene. [59]
Abundant compression floras dominated by angiosperm leaves are described from two sites of probable Pliocene age in Brunei by Wilf et al. (2022), who interpret these floras as evidence of dipterocarp-dominated lowland rainforests in the Malay Archipelago before the Pleistocene.[60]
A study on the impact of the extinct Neotropical megafauna on the variability in plant functional traits and biome geography in Central and South America is published by Dantas & Pausas (2022).[61]
A study on plant material from rock overhangs from mid-late Holocene sites along the Kawarau-Cromwell-Roxburgh Gorges in Central Otago (New Zealand), much of which was likely transported as roosting material or consumed by moa birds, and on its implications for the knowledge of the mid-late Holocene regional vegetation of Central Otago and the knowledge of vegetation changes since mid-late Holocene, is published by Pole (2022).[62]

References

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Huang, Y.-J.; Zhu, H.; Hu, J.-J.; Jia, L.-B.; Zhou, Z.-K. (2022). "New fossil evidence from the late Pliocene of Yunnan, South China, sheds light on the distribution and diversification of Sambucus L. (Adoxaceae) in the northern low latitudes". Palaeobiodiversity and Palaeoenvironments (in press). doi:10.1007/s12549-021-00519-7. S2CID 245856680.
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Hamersma, A.; Herrera, F.; Wurdack, K.; Manchester, S. R. (2022). "Belenocarpa tertiara (Berry) gen. et comb. nov. (Euphorbiaceae): Fossil Fruits with Carunculate Seeds from the Oligocene of Peru". International Journal of Plant Sciences. 183 (4): 296–306. doi:10.1086/718830. S2CID 247214597.
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Bansal, M.; Morley, R. J.; Nagaraju, S. K.; Dutta, S.; Mishra, A. K.; Selveraj, J.; Kumar, S.; Niyolia, D.; Harish, S. M.; Abdelrahim, O. B.; Hasan, S. E.; Ramesh, B. R.; Dayanandan, S.; Morley, H. P.; Ashton, P. S.; Prasad, V. (2022). "Southeast Asian Dipterocarp origin and diversification driven by Africa-India floristic interchange". Science. 375 (6579): 455–460. Bibcode:2022Sci...375..455B. doi:10.1126/science.abk2177. PMID 35084986. S2CID 246360938.
Tang, K. K.; Smith, S. Y.; Atkinson, B. A. (2022). "Extending beyond Gondwana: Cretaceous Cunoniaceae from western North America". New Phytologist. 234 (2): 704–718. doi:10.1111/nph.17976. PMID 35043416.
Shi, C.; Wang, S.; Cai, H.; Zhang, H.; Long, X.; Tihelka, E.; Song, W.; Feng, Q.; Jiang, R.; Cai, C.; Lombard, N.; Li, X.; Yuan, J.; Zhu, J.; Yang, H.; Liu, X.; Xiang, Q.; Zhao, Z.; Long, C.; Schneider, H.; Zhang, X.; Peng, H.; Li, D.-Z.; Fan, Y.; Engel, M. S.; Wang, Y.; Spicer, R. A. (2022). "Fire-prone Rhamnaceae with South African affinities in Cretaceous Myanmar amber". Nature Plants. 8 (2): 125–135. doi:10.1038/s41477-021-01091-w. PMID 35102275. S2CID 246443363.
El-Noamani, Z. M. (2022). "Ficoxylon fusiforme (Moraceae), a New Species from Upper Cretaceous Nubian Sandstone in Southern Egypt". Egyptian Journal of Botany. 62 (1): 31–44. doi:10.21608/ejbo.2021.53712.1594. S2CID 235547996.
Dong, J.-L.; Li, Z.; Gao, J.-X.; Sun, B.-N.; He, Y.-L. (2022). "Ficus leaves within the Ficus subgenus Urostigma (Moraceae) from the middle Miocene in South China and their biogeography implications". Review of Palaeobotany and Palynology: Article 104671. doi:10.1016/j.revpalbo.2022.104671.
Manchester, S. R.; Judd, W. S. (2022). "Extinct Anacardiaceous Samaras and Sumac-Like Leaves from the Eocene of Western North America". International Journal of Plant Sciences. doi:10.1086/719948.
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Heřmanová, Z.; Čepičková, J.; Kvaček, J.; von Balthazar, M.; Schönenberger, J. (2022). "A multicarpellate fruit from Late Cretaceous sediments of South Bohemia, Czech Republic". Palaeontologia Electronica. 25 (1): Article number 25.1.5A. doi:10.26879/1192.
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Palazzesi, L.; Hidalgo, O.; Barreda, V. D.; Forest, F.; Höhna, S. (2022). "The rise of grasslands is linked to atmospheric CO₂ decline in the late Palaeogene". Nature Communications. 13 (1): Article number 293. Bibcode:2022NatCo..13..293P. doi:10.1038/s41467-021-27897-y. PMC 8755714. PMID 35022396.
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Li, Y.; Wang, Y.-D.; Feldberg, K.; Wang, S.; Shi, C.; Cui, Y.-M.; Zhang, X.-Q. (2022). "New insights into the moss genus Vetiplanaxis with a description of V. obtusus sp. nov. from the mid-Cretaceous Kachin amber, Myanmar". Review of Palaeobotany and Palynology. 301 (in press): Article 104659. doi:10.1016/j.revpalbo.2022.104659. S2CID 247925481.
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[1] Deng, S.; Lu, Y.; Fan, R.; Luo, Z.; Ma, X.; Lü, D.; Sun, Y. (2022). "Lycopsid Annalepis Fliche from the Middle Triassic of the Ordos Basin, North China and reviews of the genus". Review of Palaeobotany and Palynology. in press: Article 104660. doi:10.1016/j.revpalbo.2022.104660. S2CID 248006937.

[2] Herrera, F.; Testo, W. L.; Field, A. R.; Clark, E. G.; Herendeen, P. S.; Crane, P. R.; Shi, G. (2022). "A permineralized Early Cretaceous lycopsid from China and the evolution of crown clubmosses". New Phytologist. 233 (5): 2310–2322. doi:10.1111/nph.17874. PMID 34981832. S2CID 245670357.

[3] Li, Y.; Li, Y.-D.; Wang, Y.-D.; Schneider, H.; Shi, G.-L. (2022). "Re-appraisal of lacewing mimicry of liverworts from the mid-Cretaceous Kachin amber, Myanmar with a description of Selaginella cretacea sp. nov. (Selaginellales, Selaginellaceae)". Cretaceous Research. 133: Article 105143. doi:10.1016/j.cretres.2022.105143. S2CID 245939368.

[4] Trevisan, C.; Dutra, T.; Ianuzzi, R.; Sander, A.; Wilberger, T.; Manríquez, L.; Mansilla, H.; Leppe, M. (2022). "Coniopteris antarctica sp. nov. (Pteridophyta) and associated plant assemblage from the Upper Cretaceous of Rip Point, Nelson Island, Antarctica". Cretaceous Research (in press): Article 105185. doi:10.1016/j.cretres.2022.105185. S2CID 247684239.

[5] Pšenička, J.; Zhou, W.; Boyce, C. K.; Votočková Frojdová, J.; Bek, J.; Opluštil, S.; Wang, J. (2022). "Two new leptosporangiate ferns from in situ volcanic ash of the Whetstone Horizon (Kladno Formation, Pennsylvanian), Pilsen Basin, Czech Republic". Review of Palaeobotany and Palynology. 299: Article 104608. doi:10.1016/j.revpalbo.2022.104608.

[6] Ren, W.-X.; Wu, G.-T.; Han, L.; Hua, Y.-F.; Sun, B.-N. (2022). "New species of fossil Dryopterites from the Lower Cretaceous in the Zhongkouzi Basin, Beishan area, Northwest China, and its geological significance". Historical Biology: An International Journal of Paleobiology (in press): 1–8. doi:10.1080/08912963.2021.2022135. S2CID 245694205.

[7] Barbosa, C.; Correia, P.; Muchagata, J.; Domingos, R.; Sá, A. A. (2022). "Phyllotheca douroensis sp. nov., a new equisetalean fossil-species from the Douro Carboniferous Basin (Upper Pennsylvanian; NW Portugal): palaeobiogeographical, systematic and evolutionary implications". Biosis: Biological Systems. 3 (1): e001. doi:10.37819/biosis.003.01.0162.

[8] Mendes, M. M.; Kvaček, J. (2022). "Frenelopsis antunesii sp. nov., a new cheirolepidiaceous conifer from the Lower Cretaceous of Figueira da Foz Formation in western Portugal". Review of Palaeobotany and Palynology. 300 (in press): Article 104643. doi:10.1016/j.revpalbo.2022.104643. S2CID 247433809.

[9] Sadowski, E.-M.; Schmidt, A. R.; Kunzmann, L. (2022). "The hyperdiverse conifer flora of the Baltic amber forest". Palaeontographica Abteilung B. doi:10.1127/palb/2022/0078.

[10] Andruchow-Colombo, A.; Gandolfo, M. A.; Escapa, I. H.; Cúneo, N. R. (2022). "New genus of Cupressaceae from the Upper Cretaceous of Patagonia (Argentina) fills a gap in the evolution of the ovuliferous complex in the family". Journal of Systematics and Evolution (in press). doi:10.1111/jse.12842. S2CID 247335891.

[Aciphyllum-11] Barbacka, M.; Górecki, A.; Pacyna, G.; Pieńkowski, G.; Philippe, M.; Bóka, K.; Ziaja, J.; Jarzynka, A.; Qvarnström, M.; Niedźwiedzki, G. (2022). "Early Jurassic coprolites: insights into palaeobotany and the feeding behaviour of dinosaurs". Papers in Palaeontology. 8 (2): e1425. doi:10.1002/spp2.1425. S2CID 247688865.

[12] Berry, K. (2022). "Conifer turnover across the K/Pg boundary in Colorado, U.S.A., parallels South American patterns: New and emerging perspectives". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 303 (1): 11–28. doi:10.1127/njgpa/2022/1035. S2CID 246455719.

[13] Maccracken, S. A.; Miller, I. M.; Johnson, K. R.; Sertich, J. M.; Labandeira, C. C. (2022). "Insect herbivory on Catula gettyi gen. et sp. nov. (Lauraceae) from the Kaiparowits Formation (Late Cretaceous, Utah, USA)". PLOS ONE. 17 (1): e0261397. Bibcode:2022PLoSO..1761397M. doi:10.1371/journal.pone.0261397. PMC 8782542. PMID 35061696.

[TJES312-14] Akkemik, Ü.; Iamandei, S.; Çelik, H. (2022). "Further contribution to the early Miocene woody flora of Galatian Volcanic Province from Doğanyurt Village, Ankara (Turkey)". Turkish Journal of Earth Sciences. 31 (2): 208–234. doi:10.3906/yer-2103-7 (inactive 2022-04-07).{{cite journal}}: CS1 maint: DOI inactive as of April 2022 (link)

[15] Rozefelds, A. C.; Rudall, P. J.; Herne, M. C.; Milroy, A. K.; Bridgeman, J. (2022). "A Fossil Syncarpous Fruit from Australia Provides Support for a Gondwanan History for the Screw Pines (Pandanus, Pandanaceae)". International Journal of Plant Sciences. 183 (4): 320–329. doi:10.1086/719431. S2CID 247378720.

[PDKumaretal-16] Kumar, S.; Hazra, T.; Spicer, R. A.; Hazra, M.; Spicer, T. E. V.; Bera, S.; Khan, M. A. (2022). "Coryphoid palms from the K-Pg boundary of central India and their biogeographical implications: Evidence from megafossil remains". Plant Diversity (in press). doi:10.1016/j.pld.2022.01.001. S2CID 246084708.

[17] Greenwood, D. R.; Conran, J. G.; West, C. K. (2022). "Palm fronds from western Canada are the northernmost palms from the Late Cretaceous of North America and may include the oldest Arecaceae". Review of Palaeobotany and Palynology. 301 (in press): Article 104641. doi:10.1016/j.revpalbo.2022.104641. S2CID 247343236.

[18] Onstein, R. E.; Kissling, W. D.; Linder, H. P. (2022). "The megaherbivore gap after the non-avian dinosaur extinctions modified trait evolution and diversification of tropical palms". Proceedings of the Royal Society B: Biological Sciences. 289 (1972): Article ID 20212633. doi:10.1098/rspb.2021.2633. PMID 35414237. S2CID 248119885.

[Langeria2022-19] Huegele, I. B.; Manchester, S. R. (2022). "Newly Recognized Reproductive Structures Linked with Langeria from the Eocene of Washington, USA, and their Affinities with Platanaceae". International Journal of Plant Sciences (in press). doi:10.1086/720138. S2CID 247907696.

[20] Moiseeva, M. G.; Kodrul, T. M.; Tekleva, M. V.; Maslova, N. P.; Wu, X.; Jin, J. (2022). "Fossil Leaves of Meliosma (Sabiaceae) With Associated Pollen and a Eupodid Mite From the Eocene of Maoming Basin, South China". Frontiers in Ecology and Evolution. 9: Article 770687. doi:10.3389/fevo.2021.770687.

[21] Dong, J.-L.; Gao, J.-X.; Li, Z.; Sun, B.-N. (2022). "A tropical lotus from the middle Miocene tropical rainforest flora of South China". Review of Palaeobotany and Palynology. 299: Article 104611. doi:10.1016/j.revpalbo.2022.104611. S2CID 246467568.

[Huegele2021-22] Huegele, I.B.; Zhu, H.; Zhao, B.; Wang, Y.-F.; Manchester, S. R. (2021). "Trans-Beringial Distribution of Platimeliphyllum (Platanaceae) in the Eocene of Eastern Asia and Western North America". International Journal of Plant Sciences. 183 (2): 139–153. doi:10.1086/717692. S2CID 239529168.

[23] Gobo, W. V.; Kunzmann, L.; Iannuzzi, R.; Bachelier, J. B.; Coiffard, C. (2022). "First evidence of ranunculids in Early Cretaceous tropics". Scientific Reports. 12 (1): Article number 5040. Bibcode:2022NatSR..12.5040G. doi:10.1038/s41598-022-07920-y. PMC 8943169. PMID 35322034.

[Phytotaxa5312-24] Doweld, A. B. (2022). "New names of Ilex and Ilexpollenites (Aquifoliaceae), extant and fossil: Addendum Notulae Systematicae ad Palaeofloram Europaeam spectantes II. Aquifoliaceae". Phytotaxa. 531 (2): 143–146. doi:10.11646/phytotaxa.531.2.7.

[25] Huang, Y.-J.; Zhu, H.; Hu, J.-J.; Jia, L.-B.; Zhou, Z.-K. (2022). "New fossil evidence from the late Pliocene of Yunnan, South China, sheds light on the distribution and diversification of Sambucus L. (Adoxaceae) in the northern low latitudes". Palaeobiodiversity and Palaeoenvironments (in press). doi:10.1007/s12549-021-00519-7. S2CID 245856680.

[Paleobowdichia-26] Herendeen, P. S.; Cardoso, D. B. O. S.; Herrera, F.; Wing, S. L. (2022). "Fossil papilionoids of the Bowdichia clade (Leguminosae) from the Paleogene of North America". American Journal of Botany. 109 (1): 130–150. doi:10.1002/ajb2.1808. PMID 35014023. S2CID 245897924.

[27] Jia, L.-B.; Hu, J.-J.; Zhang, S.-T.; Su, T.; Spicer, R. A.; Liu, J.; Yang, J.-C.; Zou, P.; Huang, Y.-J.; Zhou, Z.-K. (2022). "Bauhinia (Leguminosae) Fossils from the Paleogene of Southwestern China and Its Species Accumulation in Asia". Diversity. 14 (3): Article 173. doi:10.3390/d14030173.

[28] Hamersma, A.; Herrera, F.; Wurdack, K.; Manchester, S. R. (2022). "Belenocarpa tertiara (Berry) gen. et comb. nov. (Euphorbiaceae): Fossil Fruits with Carunculate Seeds from the Oligocene of Peru". International Journal of Plant Sciences. 183 (4): 296–306. doi:10.1086/718830. S2CID 247214597.

[29] Hazra, T.; Mahato, S.; Bera, S.; Khan, M. A. (2022). "First fossil evidence of Indian tulip tree". Botany Letters (in press): 1–10. doi:10.1080/23818107.2021.2014358. S2CID 246435429.

[30] Bansal, M.; Morley, R. J.; Nagaraju, S. K.; Dutta, S.; Mishra, A. K.; Selveraj, J.; Kumar, S.; Niyolia, D.; Harish, S. M.; Abdelrahim, O. B.; Hasan, S. E.; Ramesh, B. R.; Dayanandan, S.; Morley, H. P.; Ashton, P. S.; Prasad, V. (2022). "Southeast Asian Dipterocarp origin and diversification driven by Africa-India floristic interchange". Science. 375 (6579): 455–460. Bibcode:2022Sci...375..455B. doi:10.1126/science.abk2177. PMID 35084986. S2CID 246360938.

[Tang2022-31] Tang, K. K.; Smith, S. Y.; Atkinson, B. A. (2022). "Extending beyond Gondwana: Cretaceous Cunoniaceae from western North America". New Phytologist. 234 (2): 704–718. doi:10.1111/nph.17976. PMID 35043416.

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