2022 in reptile paleontology

This list of fossil reptiles described in 2022 is a list of new taxa of fossil reptiles that were described during the year 2022, as well as other significant discoveries and events related to reptile paleontology that occurred in 2022.

List of years in reptile paleontology
In archosaur paleontology
2019
2020
2021
2022
2023
2024
2025
In paleontology
2019
2020
2021
2022
2023
2024
2025

Squamates

New taxa
Name Novelty Status Authors Age Type locality Country Notes Images

Abronia cuyama[1]

Sp. nov

Valid

Scarpetta & Ledesma

Miocene

Caliente Formation

 United States
( California)

A species of Abronia.

Archaeovaranus[2]

Gen. et sp. nov

Dong et al.

Early Eocene

Yuhuangding Formation

 China

A stem-varanid. The type species is A. lii.

Blutwurstia[3]

Gen. et sp. nov

Valid

Smith, Bhullar & Bloch

Early Eocene

Willwood Formation

 United States
( Wyoming)

A member of Pan-Xenosaurus. The type species is B. oliviae.

Psammophis odysseus[4]

 Sp. nov In press Szyndlar & Georgalis Late Miocene  Spain A species of Psammophis.

Retinosaurus[5]

Gen. et sp. nov

Valid

Čerňanský et al.

Early Cretaceous (Albian)

Hkamti amber

 Myanmar

A scincomorph lizard, possibly a member of Pan-Xantusiidae. The type species is R. hkamtiensis.

Research
  • A study aiming to determine whether the squamate fossil record contains reliable phylogenetic information in spite of its incompleteness is published by Woolley et al. (2022).[6]
  • A vertebra of a monitor lizard, representing the first published record of a non-snake squamate from the Neogene of Pakistan (probably Miocene Chinji Formation), is described by Villa & Delfino (2022), who evaluate the implications of this finding for the palaeoenvironmental reconstructions of the Siwaliks during the Miocene.[7]
  • A study on the diversification of feeding and locomotory strategies of mosasauroids is published by Cross et al. (2022).[8]
  • The first occurrences of Mosasaurus hoffmannii are reported from the Ouled Abdoun Basin (Morocco) by Rempert et al. (2022), extending the known range of this species.[9]
  • El-Hares et al. (2022) describe new fossil material of colubroidean snakes from the Eocene Birket Qarun Locality 2 (Egypt), including a vertebra of a member or a relative of the genus Procerophis (otherwise known from the early Eocene of India) and the first known caudal vertebrae of Renenutet enmerwer, as well as a vertebra representing the first record of an amphisbaenian from the Paleogene of Egypt reported to date.[10]

Ichthyosauromorphs

New Taxa
Name Novelty Status Authors Age Type Locality Location Notes Images

Baisesaurus[11]

Gen. et sp. nov

Valid

Ren et al.

Early Triassic (Olenekian)

Luolou Formation

 China

An early member of Ichthyosauromorpha . The type species is B. robustus
Magnipterygius[12] Gen. et sp. nov Valid Maisch & Matzke Early Jurassic Posidonia Shale  Germany A stenopterygiid ichthyosaur. The type species is M. huenei

Research
  • Fossil material of ichthyopterygians, including a limb bone (probably a humerus) representing one of the largest specimens of early Spathian marine reptiles known to date, is described from the Lower Triassic Zhitkov Formation (Russky Island, Primorsky Krai, Russia) by Nakajima et al. (2022).[13]
  • Fossil material of giant ichthyosaurs is described from the Upper Triassic Kössen Formation (Switzerland) by Sander et al. (2022), who evaluate the implications of the studied fossils for the knowledge of the global distribution and ecological diversity of giant Norian and Rhaetian ichthyosaurs.[14]
  • A study on ichthyosaur vertebral centra from the Upper Jurassic Bernbjerg Formation (Greenland), aiming to determine whether vertebral ratios can be used to assign disarticulated and possibly weathered centra to a region in the vertebral column of ichthyosaurs, is published by Holm, Delsett & Alsen (2022).[15]
  • Fossil material of ichthyosaurs is described from the Valanginian and Hauterivian of Austria by Lukeneder et al. (2022), who interpret the studied fossils as representing two distinct taxa, probably with different feeding ecologies, and evaluate the implications of these fossils for the knowledge of the diversity of ichthyosaurs in the Early Cretaceous.[16]
  • Roberts, Engelschiøn & Hurum (2022) describe a new specimen of Phalarodon fraasi from the Ladinian Blanknuten Member of the Botneheia Formation (Svalbard, Norway), representing the first three-dimensional mixosaurid skull recovered from this formation.[17]
  • A study on the anatomy and phylogenetic relationships of "Ichthyosaurus" zetlandicus is published by Laboury et al. (2022), who transfer this species to the genus Temnodontosaurus.[18]
  • A study on the pre- and postnatal ontogenetic changes in the skull of Stenopterygius quadriscissus is published by Miedema & Maxwell (2022).[19]
  • Two ichthyosaur specimens (a nearly complete skeleton of a member of the genus Aegirosaurus and an isolated tail of an indeterminate ichthyosaur) preserved with soft tissue are described from the Upper Jurassic (Tithonian) Eichstätt Plattenkalk (Germany) by Delsett et al. (2022).[20]

Sauropterygians
  • Description of two placodont dentaries from the Muschelkalk of the Netherlands and Germany, possibly belonging to a member of the genus Placodus belonging or related to the species P. gigas, and a study on the implications of these fossils for the knowledge of the ontogenetic changes in placodont dentaries is published by Klein et al. (2022).[21]
  • Redescription of the type material of Ischyrodon meriani is published by Madzia, Sachs & Klug (2022).[22]
  • A study on the diversity and ontogenetic changes of cervical vertebral shapes in elasmosaurids is published by Brum et al. (2022).[23]

Turtles

New taxa
Name Novelty Status Authors Age Type locality Location Notes Images

Dortoka vremiri[24]

Sp. nov

Valid

Augustin et al.

Late Cretaceous (Maastrichtian)

Sînpetru Formation

 Romania

A stem-pleurodiran belonging the family Dortokidae.

Hutchemys walkerorum[25]

Sp. nov

In press

Jasinski et al.

Late Cretaceous (Maastrichtian)

Hell Creek Formation

 United States
( North Dakota)

A member of Pan-Trionychidae belonging to the family Plastomenidae.
Jimemys[26] Gen. et sp. nov In press Edgar et al. Late Cretaceous (Santonian-Campanian) Milk River Formation  Canada

A member of Pan-Trionychidae belonging to the family Plastomenidae. The type species is J. glaebosus.

Testudo lohanica[27]

Sp. nov

Pérez-García et al.

Late Miocene

 Romania

A tortoise, a species of Testudo.

Research
  • Review of the development and evolutionary history of the scute patterns of the carapace of extant and fossil turtles is published by Ascarrunz & Sánchez-Villagra (2022).[28]
  • A clutch of turtle eggs with unique attributes of the eggshell is described from the Upper Cretaceous Kaiparowits Formation (Utah, United States) by Ferguson & Tapanila (2022), who name a new oospecies Testudoolithus tuberi.[29]
  • Description of the anatomy of the skull of Lakotemys australodakotensis is published by Rollot et al. (2022).[30]
  • Danilov et al. (2022) describe fossil material of Campanian pan-chelonioid turtles from the Beloe Ozero locality (Rybushka Formation; Saratov Oblast, Russia), including specimens of Protostega gigas with estimated size corresponding to those of the largest specimens from North America, representing the first record of this species outside North America reported to date.[31]

Archosaurs
Main article: 2022 in archosaur paleontology

Other reptiles

New taxa
Name Novelty Status Authors Age Type locality Country Notes Images

Champsosaurus norelli[32]

Sp. nov

Valid

Brownstein

Paleocene (Tiffanian)

Polecat Bench Formation

 United States
( Wyoming)

Kosmodraco[32]

Gen. et comb. et sp. nov

Valid

Brownstein

Paleocene

 United States
( North Dakota
 Wyoming)

A member of Choristodera belonging to the family Simoedosauridae. The type species is "Simoedosaurus" dakotensis Erickson (1987); genus also includes new species K. magnicornis.

Navajosphenodon[33]

Gen. et sp. nov

Valid

Simões, Kinney-Broderick & Pierce

Early Jurassic (Sinemurian-Pliensbachian)

Kayenta Formation

 United States
( Arizona)

A rhynchocephalian belonging to the group Sphenodontinae. The type species is N. sani.

Quasicolognathus[34]

Gen. et sp. nov

Valid

Sues, Kligman & Schoch

Middle Triassic (Ladinian)

Erfurt Formation

 Germany

A reptile with similarities to Colognathus, belonging to the new family Colognathidae. The type species is Q. eothen.

Research
  • A study on the proportions of skull and limb bones in mesosaur specimens, aiming to determine whether there are statistically significant morphological differences through ontogenetic development among mesosaurs coming from Africa, Brazil and Uruguay, and evaluating the implications of the developmental pattern found in mesosaurs for the knowledge of the phylogeny and evolution of growth patterns of early amniotes, is published by Núñez Demarco, Ferigolo & Piñeiro (2022).[35]
  • A study on tooth implantation, replacement and attachment in Scoloparia glyphanodon is published by Jenkins & Bhullar (2022).[36]
  • Revision and a study on the phylogenetic affinities of bradysaurian pareiasaurs from the Guadalupian of the Karoo Basin (South Africa) is published by Van den Brandt et al. (2022).[37]
  • A study on the anatomy of the skull of Captorhinus aguti, indicative of the presence of distinct anatomical modules on each side of the skull, and evaluating the implications of this modularity pattern for the knowledge of the evolution and function of temporal openings in amniote skulls, is published by Werneburg & Abel (2022).[38]
  • New fossil material of Moradisaurus grandis (two partial skulls of juvenile individuals) is described from the Permian Moradi Formation (Niger) by Sidor et al. (2022).[39]
  • A large rib bearing an osteoderm is described from the Upper Triassic Kössen Formation (Switzerland) by Scheyer et al. (2022), who interpret this specimen as a member or a relative of the family Saurosphargidae with potential affinities to the genus Largocephalosaurus, potentially extending the occurrence of saurosphargids about 35 million years into the Late Triassic.[40]
  • Description of the neomorphic ossification between the parietal, quadrate and squamosal in the skulls of Coeruleodraco jurassicus and Philydrosaurus proseilus is published by Qin, Yi & Gao (2022).[41]
  • Description of new specimens of Hyperodapedon from Brazil with apically serrated teeth crowns, representing new maxillary tooth morphotype and the first records of serrated teeth in rhynchosaurs, and a study aiming to determine whether the maxillary crown morphology of rhynchosaurs is taxonomically informative or reflects ontogeny, is published by Scartezini & Soares (2022).[42]
  • A study on the anatomy and phylogenetic affinities of Tricuspisaurus thomasi and Variodens inopinatus is published by Chambi-Trowell et al. (2022).[43]
  • A study on the skeletal anatomy and phylogenetic affinities of Shringasaurus indicus is published by Sengupta & Bandyopadhyay (2022).[44]

Reptiles in general
  • A study on the impact of the body size and shape on drag in ichthyosaurs and plesiosaurs, and on the impact of this relationship on the evolution of trunk length and neck proportions in Sauropterygia, is published by Gutarra et al. (2022).[45]
  • Tracks produced by squamates or rhynchocephalians are described from the Lower Cretaceous Botucatu Formation (Brazil) by Buck et al. (2022), representing the first known evidence of the presence of lepidosaurs in the ancient Botucatu desert.[46]

References
  1. Scarpetta, S. G.; Ledesma, D. T. (2022). "A strikingly ornamented fossil alligator lizard (Squamata: Abronia) from the Miocene of California". Zoological Journal of the Linnean Society. doi:10.1093/zoolinnean/zlac024.
  2. Dong, L.; Wang, Y.-Q.; Zhao, Q.; Vasilyan, D.; Wang, Y.; Evans, S. E. (2022). "A new stem-varanid lizard (Reptilia, Squamata) from the early Eocene of China". Philosophical Transactions of the Royal Society B: Biological Sciences. 377 (1847): Article ID 20210041. doi:10.1098/rstb.2021.0041. PMC 8819366. PMID 35125002.
  3. Smith, K. T.; Bhullar, B.-A. S.; Bloch, J. I. (2022). "New diminutive Eocene lizard reveals high K-Pg survivorship and taxonomic diversity of stem xenosaurs in North America". American Museum Novitates (3986): 1–36. doi:10.1206/3986.1. hdl:2246/7293.
  4. Georgalis, G. L.; Szyndlar, Z. "First occurrence of Psammophis (Serpentes) from Europe witnesses another Messinian herpetofaunal dispersal from Africa – biogeographic implications and a discussion of the vertebral morphology of psammophiid snakes". The Anatomical Record. In press. doi:10.1002/ar.24892. PMID 35139258.
  5. Čerňanský, A.; Stanley, E. L.; Daza, J. D.; Bolet, A.; Arias, J. S.; Bauer, A. M.; Vidal-García, M.; Bevitt, J. J.; Peretti, A. M.; Aung, N. N.; Evans, S. E. (2022). "A new Early Cretaceous lizard in Myanmar amber with exceptionally preserved integument". Scientific Reports. 12 (1): Article number 1660. doi:10.1038/s41598-022-05735-5. PMC 8803969. PMID 35102237.
  6. Woolley, C. H.; Thompson, J. R.; Wu, Y.-H.; Bottjer, D. J.; Smith, N. D. (2022). "A biased fossil record can preserve reliable phylogenetic signal". Paleobiology. in press. doi:10.1017/pab.2021.45.
  7. Villa, A.; Delfino, M. (2022). "First fossil of Varanus Merrem, 1820 (Squamata: Varanidae) from the Miocene Siwaliks of Pakistan". Geodiversitas. 44 (7): 229–235. doi:10.5252/geodiversitas2022v44a7.
  8. Cross, S. R. R.; Moon, B. C.; Stubbs, T. L.; Rayfield, E. J.; Benton, M. J. (2022). "Climate, competition, and the rise of mosasauroid ecomorphological disparity". Palaeontology. 65 (2): e12590. doi:10.1111/pala.12590.
  9. Rempert, Trevor; Vinkeles Melchers, Alexander P.M.; Rempert, Ashley N.; Haque, Muhammad R.; Armstrong, Andrew. "Occurrence of Mosasaurus hoffmannii Mantell, 1829 (Squamata, Mosasauridae) in the Maastrichtian Phosphates of Morocco". Journal of Paleontological Sciences.
  10. El-Hares, M. A.; Zaher, H.; El-Mekkawy, D.; El-Sayed, S.; Seiffert, E. R.; Sallam, H. M. (2022). "New records of legless squamates from the lowest upper Eocene deposits of the Fayum Depression, Egypt". Journal of Vertebrate Paleontology. in press: e1992770. doi:10.1080/02724634.2021.1992770.
  11. Ren, J.; Jiang, H.; Xiang, K.; Sullivan, C.; He, Y.; Cheng, L.; Han, F. (2022). "A new basal ichthyosauromorph from the Lower Triassic (Olenekian) of Zhebao, Guangxi Autonomous Region, South China". PeerJ. 10: e13209. doi:10.7717/peerj.13209.
  12. Maisch, Michael; Matzke, Andreas. "Magnipterygius huenei n. gen. n. sp., a new small stenopterygiid (Reptilia: Ichthyosauria) from the Posidonienschiefer Formation of SW Germany". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 303 (2): 169–201. doi:10.1127/njgpa/2022/1042.
  13. Nakajima, Y.; Shigeta, Y.; Houssaye, A.; Zakharov, Y. D.; Popov, A. M.; Sander, P. M. (2022). "Early Triassic ichthyopterygian fossils from the Russian Far East". Scientific Reports. 12 (1): Article number 5546. doi:10.1038/s41598-022-09481-6.
  14. Sander, P. M.; Romero Pérez de Villar, P.; Furrer, H.; Wintrich, T. (2022). "Giant Late Triassic ichthyosaurs from the Kössen Formation of the Swiss Alps and their paleobiological implications". Journal of Vertebrate Paleontology: e2046017. doi:10.1080/02724634.2021.2046017.
  15. Holm, T. B.; Delsett, L. L.; Alsen, P. (2022). "Vertebral size ratios and the ichthyosaurian vertebral column – a case study based on Late Jurassic fossils from North-East Greenland". Bulletin of the Geological Society of Denmark. 70: 1–17. doi:10.37570/bgsd-2022-70-01.
  16. Lukeneder, A.; Zverkov, N.; Kaurin, C.; Blüml, V. (2022). "First Early Cretaceous ichthyosaurs of Austria and the problem of Jurassic–Cretaceous ichthyosaurian faunal turnover". Cretaceous Research. in press: Article 105224. doi:10.1016/j.cretres.2022.105224.
  17. Roberts, A. J.; Engelschiøn, V. S.; Hurum, J. H. (2022). "First three-dimensional skull of the Middle Triassic mixosaurid ichthyosaur Phalarodon fraasi from Svalbard, Norway". Acta Palaeontologica Polonica. 67 (1): 51–62. doi:10.4202/app.00915.2021.
  18. Laboury, A.; Bennion, R. F.; Thuy, B.; Weis, R.; Fischer, V. (2022). "Anatomy and phylogenetic relationships of Temnodontosaurus zetlandicus (Reptilia: Ichthyosauria)". Zoological Journal of the Linnean Society. in press. doi:10.1093/zoolinnean/zlab118.
  19. Miedema, F.; Maxwell, E. E. (2022). "Ontogenetic variation in the skull of Stenopterygius quadriscissus with an emphasis on prenatal development". Scientific Reports. 12 (1): Article number 1707. doi:10.1038/s41598-022-05540-0. PMC 8807662. PMID 35105895.
  20. Delsett, L. L.; Friis, H.; Kölbl-Ebert, M.; Hurum, J. H. (2022). "The soft tissue and skeletal anatomy of two Late Jurassic ichthyosaur specimens from the Solnhofen archipelago". PeerJ. 10: e13173. doi:10.7717/peerj.13173.
  21. Klein, N.; Wintrich, T.; Hagdorn, H.; Spiller, D.; Winkelhorst, H.; Goris, G.; Scheyer, T. M. (2022). "Placodus (Placodontia, Sauropterygia) dentaries from Winterswijk, The Netherlands (middle Anisian) and Hünfeld, Hesse, Germany (late Anisian) with comments on ontogenetic changes". PalZ. in press. doi:10.1007/s12542-022-00614-w.
  22. Madzia, D.; Sachs, S.; Klug, C. (2022). "Historical significance and taxonomic status of Ischyrodon meriani (Pliosauridae) from the Middle Jurassic of Switzerland". PeerJ. 10: e13244. doi:10.7717/peerj.13244.
  23. Brum, A. S.; Simões, T. R.; Souza, G. A.; Pinheiro, A. E. P.; Figueiredo, R. G.; Caldwell, M. W.; Sayão, J. M.; Kellner, A. W. A. (2022). "Ontogeny and evolution of the elasmosaurid neck highlight greater diversity of Antarctic plesiosaurians". Palaeontology. 65 (2): e12593. doi:10.1111/pala.12593.
  24. Augustin, F. J.; Csiki-Sava, Z.; Matzke, A. T.; Botfalvai, G.; Rabi, M. (2022). "A new latest Cretaceous pleurodiran turtle (Testudinata: Dortokidae) from the Haţeg Basin (Romania) documents end-Cretaceous faunal provinciality and selective survival during the K-Pg extinction". Journal of Systematic Palaeontology. 19 (15): 1059–1081. doi:10.1080/14772019.2021.2009583.
  25. Jasinski, S. E.; Heckert, A. B.; Sailar, C.; Lichtig, A. J.; Lucas, S. G.; Dodson, P. (2022). "A softshell turtle (Testudines: Trionychidae: Plastomeninae) from the uppermost Cretaceous (Maastrichtian) Hell Creek formation, North Dakota, USA, with implications for the evolutionary relationships of plastomenines and other trionychids". Cretaceous Research. in press: Article 105172. doi:10.1016/j.cretres.2022.105172.
  26. Edgar, Shauna C.; Brinkman, Don B.; Ryan, Michael J.; Evans, David C. (2022-03-11). "A new plastomenid trionychid (Testudines: Pan-Trionychidae) from Milk River Formation of southern Alberta (Cretaceous: Santonian)". Canadian Journal of Earth Sciences. doi:10.1139/cjes-2021-0040.
  27. Pérez-García, A.; Martín-Jiménez, M.; Vlachos, E.; Codrea, V. (2022). "The most complete extinct species of Testudo (Testudines, Testudinidae) defined by several well-preserved skeletons from the late Miocene of Romania". Journal of Systematic Palaeontology. in press. doi:10.1080/14772019.2022.2028025.
  28. Ascarrunz, E.; Sánchez-Villagra, M. R. (2022). "The macroevolutionary and developmental evolution of the turtle carapacial scutes". Vertebrate Zoology. 72: 29–46. doi:10.3897/vz.72.e76256.
  29. Ferguson, A. L.; Tapanila, L. (2022). "Rare clutch of Cretaceous turtle eggs preserved in the Kaiparowits Formation of southern Utah". Cretaceous Research. in press: Article 105197. doi:10.1016/j.cretres.2022.105197.
  30. Rollot, Y.; Evers, S. W.; Cifelli, R. L.; Joyce, W. G. (2022). "New insights into the cranial osteology of the Early Cretaceous paracryptodiran turtle Lakotemys australodakotensis". PeerJ. 10: e13230. doi:10.7717/peerj.13230.
  31. Danilov, I. G.; Obraztsova, E. M.; Arkhangelsky, M. S.; Ivanov, A. V.; Averianov, A. O. (2022). "Protostega gigas and other sea turtles from the Campanian of Eastern Europe, Russia". Cretaceous Research. in press: Article 105196. doi:10.1016/j.cretres.2022.105196.
  32. Brownstein, C. D. (2022). "High morphological disparity in a bizarre Paleocene fauna of predatory freshwater reptiles". BMC Ecology and Evolution. 22 (1): Article number 34. doi:10.1186/s12862-022-01985-z. PMID 35313822.
  33. Simões, T. R.; Kinney-Broderick, G.; Pierce, S. E. (2022). "An exceptionally preserved Sphenodon-like sphenodontian reveals deep time conservation of the tuatara skeleton and ontogeny". Communications Biology. 5 (1): Article number 195. doi:10.1038/s42003-022-03144-y. PMC 8894340. PMID 35241764.
  34. Sues, H.-D.; Kligman, B. T.; Schoch, R. R. (2022). "An unusual Colognathus-like reptile from the Middle Triassic (Ladinian) Erfurt Formation of Germany". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 303 (2): 227–238. doi:10.1127/njgpa/2022/1046.
  35. Núñez Demarco, P.; Ferigolo, J.; Piñeiro, G. (2022). "Isometry in mesosaurs: Implications for growth patterns in early amniotes". Acta Palaeontologica Polonica. in press. doi:10.4202/app.00931.2021.
  36. Jenkins, K. M.; Bhullar, B.-A. S. (2022). "Tooth Implantation and Attachment in Scoloparia glyphanodon (Parareptilia: Procolophonidae)". Bulletin of the Peabody Museum of Natural History. 63 (1): 27–30. doi:10.3374/014.063.0103.
  37. Van den Brandt, M. J.; Abdala, F.; Benoit, J.; Day, M. O.; Groenewald, D. P.; Rubidge, B. S. (2022). "Taxonomy, phylogeny and stratigraphical ranges of middle Permian pareiasaurs from the Karoo Basin of South Africa". Journal of Systematic Palaeontology. in press. doi:10.1080/14772019.2022.2035440.
  38. Werneburg, I.; Abel, P. (2022). "Modeling Skull Network Integrity at the Dawn of Amniote Diversification With Considerations on Functional Morphology and Fossil Jaw Muscle Reconstructions". Frontiers in Ecology and Evolution. 9: Article 799637. doi:10.3389/fevo.2021.799637.
  39. Sidor, C. A.; Ide, O.; Larsson, H. C. E.; O'Keefe, F. R.; Smith, R. M. H.; Steyer, J.-S.; Modesto, S. P. (2022). "The vertebrate fauna of the upper Permian of Niger—XI. Cranial material of a juvenile Moradisaurus grandis (Reptilia: Captorhinidae)". Journal of Vertebrate Paleontology. in press: e2030345. doi:10.1080/02724634.2021.2030345.
  40. Scheyer, T. M.; Oberli, U.; Klein, N.; Furrer, H. (2022). "A large osteoderm-bearing rib from the Upper Triassic Kössen Formation (Norian/Rhaetian) of eastern Switzerland". Swiss Journal of Palaeontology. 141 (1): Article 1. doi:10.1186/s13358-022-00244-4.
  41. Qin, W.; Yi, H.; Gao, K. (2022). "A neomorphic ossification connecting the braincase, squamosal, and quadrate in choristoderan reptiles: insights from µCT data". Fossil Record. 25 (1): 1–10. doi:10.3897/fr.25.79595.
  42. Scartezini, C. A.; Soares, M. B. (2022). "Assessing the diversity of hidden dental morphology in Hyperodapedontinae rhynchosaurs (Archosauromorpha, Rhynchosauria)". Historical Biology: An International Journal of Paleobiology. in press. doi:10.1080/08912963.2021.2022133.
  43. Chambi-Trowell, S. A. V.; Whiteside, D. I.; Skinner, M.; Benton, M. J.; Rayfield, E. J. (2022). "Phylogenetic relationships of the European trilophosaurids Tricuspisaurus thomasi and Variodens inopinatus". Journal of Vertebrate Paleontology. in press: e1999250. doi:10.1080/02724634.2021.1999250.
  44. Sengupta, S.; Bandyopadhyay, S. (2022). "The osteology of Shringasaurus indicus, an archosauromorph from the Middle Triassic Denwa Formation, Satpura Gondwana Basin, Central India". Journal of Vertebrate Paleontology. in press: e2010740. doi:10.1080/02724634.2021.2010740.
  45. Gutarra, S.; Stubbs, T. L.; Moon, B. C.; Palmer, C.; Benton, M. J. (2022). "Large size in aquatic tetrapods compensates for high drag caused by extreme body proportions". Communications Biology. 5 (1): Article number 380. doi:10.1038/s42003-022-03322-y. PMID 35484197.
  46. Buck, P. V.; Ghilardi, A. M.; Peixoto, B. C. P. M.; Aureliano, T.; Fernandes, M. A. (2022). "Lacertoid tracks from the Botucatu Formation (Lower Cretaceous) with different locomotor behaviors: A new trackmaker with novel paleoecological implications". Journal of South American Earth Sciences: Article 103825. doi:10.1016/j.jsames.2022.103825.
This article is issued from Wikipedia. The text is licensed under Creative Commons - Attribution - Sharealike. Additional terms may apply for the media files.