Araneomorphae

The Araneomorphae (also called the Labidognatha) are an infraorder of spiders. They are distinguishable by chelicerae (fangs) that point diagonally forward and cross in a pinching action, in contrast to the Mygalomorphae (tarantulas and their close kin), where they point straight down. Araneomorphs comprise the vast majority of living spiders.

Hippasa agelenoides  common funnel web spider

Araneomorph spiders
Temporal range:
Nephila inaurata (Nephilidae)
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Class: Arachnida
Order: Araneae
Suborder: Opisthothelae
Infraorder: Araneomorphae
Subdivisions
Diversity
95 families

Distinguishing characteristics

Most spider species are Araneomorphae, which have fangs that face towards each other, increasing the orientations they can employ during prey capture. They have fewer book lungs (when present), and the females typically live one year.

The Mygalomorphae have fangs that face towards the ground, and which are parallel to the long axis of the spider's body, thus they have only one orientation they can employ during prey capture. They have four pairs of book lungs, and the females often live many years.[1]

Spiders included

Almost all of the familiar spiders are included in the Araneomorphae group, the major exception are the Tarantulas. There are a few other Mygalomorphae species that live around homes or gardens, but they typically are relatively small and not easily noticed.

The Araneomorphae, to the contrary, include the weavers of spiral webs; the cobweb spiders that live in the corners of rooms, and between windows and screens; the crab spiders that lurk on the surfaces of flowers in gardens; the jumping spiders that are visible hunting on surfaces; the wolf spiders that carpet hunting sites in sunny spots; and the large huntsman spiders.

Systematics

In older schemes, the Araneomorphae were divided into two lineages, the Hypochilae (containing only the family Hypochilidae), and the Neocribellatae. The Neocribellatae were in turn divided into the Austrochiloidea, and the two series Haplogynae and Entelogynae, each containing several superfamilies. Molecular phylogenetic studies have shown that the Haplogyne in particular are not a monophyletic group. A 2020 study suggested the relationships among the major groups were as shown in the following cladogram.[2]

Araneomorphae

Filistatidae + Hypochilidae

Synspermiata

Leptonetidae

Austrochiloidea

Palpimanoidea

Entelegynae

This relationship was supported in a 2021 study, except for the division of Leptonetidae into two families:[3] The added yellow shading shows the former Haplogynae in the sense of Coddington (2005).[4]

Araneomorphae

Filistatidae + Hypochilidae

Synspermiata

Archoleptonetidae

Leptonetidae

Austrochiloidea

Palpimanoidea

Entelegynae

Table of families

Key
Genera1≥2≥10≥100
Species1–9≥10≥100≥1000
Araneomorphae families[note 1]
FamilyGeneraSpeciesCommon nameExample
Agelenidae781282araneomorph funnel-web spidersHobo spider
Amaurobiidae49274tangled nest spidersCallobius claustrarius
Ammoxenidae418
Anapidae58223
Anyphaenidae56563anyphaenid sac spidersYellow ghost spider
Araneidae1743128orb-weaver spidersZygiella x-notata
Archaeidae590pelican spidersMadagascarchaea gracilicollis
Arkyidae237
Austrochilidae310Tasmanian cave spider
Caponiidae18119Diploglena capensis
Cheiracanthiidae
(syn. Eutichuridae)
12351Cheiracanthium mildei
Cithaeronidae28
Clubionidae15618sac spidersClubiona trivialis
Corinnidae67779dark sac spidersCastianeira sp.
Ctenidae47525tropical wolf spidersBrazilian wandering spiders
Cyatholipidae2358
Cybaeidae19259
Cycloctenidae880
Deinopidae265net-casting spidersRufous net-casting spider
Desidae60297intertidal spidersPhryganoporus candidus
Dictynidae52464Nigma walckenaeri
Diguetidae215coneweb spiders
Drymusidae217false violin spiders
Dysderidae24564woodlouse hunter spidersWoodlouse spider
Eresidae998velvet spidersEresus sandaliatus
Filistatidae19164crevice weaversSouthern house spider
Gallieniellidae1056
Gnaphosidae[note 2]1582532flat-bellied ground spidersDrassodes cupreus
Gradungulidae716large-clawed spidersCarrai cave spider
Hahniidae23346dwarf sheet spiders
Hersiliidae16181tree trunk spidersHersilia savignyi
Homalonychidae13
Huttoniidae11Huttonia palpimanoides
Hypochilidae212lampshade spidersHypochilus thorelli
Lamponidae23192White-tailed spider
Leptonetidae21346Tooth cave spider
Linyphiidae6074566dwarf / money spidersLinyphia triangularis
Liocranidae31272liocranid sac spiders
Lycosidae1242419wolf spidersLycosa tarantula
Malkaridae1146shield spiders
Mecysmaucheniidae725
Megadictynidae22
Mimetidae12152pirate spidersOarces reticulatus
Miturgidae29130long-legged sac spiders
Mysmenidae13137spurred orb-weavers
Nesticidae16278cave cobweb spidersNesticella marapu
Nicodamidae727
Ochyroceratidae20216midget ground weaversTheotima minutissima
Oecobiidae6113disc web spidersOecobius navus
Oonopidae1141801dwarf hunting spidersOonops domesticus
Orsolobidae30188
Oxyopidae9457lynx spidersGreen lynx spider
Pacullidae438
Palpimanidae18150palp-footed spiders
Penestomidae19
Periegopidae13
Philodromidae30539philodromid crab spidersPhilodromus dispar
Pholcidae771666daddy long-legs spidersPholcus phalangioides
Phrurolithidae13205
Physoglenidae1372
Phyxelididae1464
Pimoidae441Pimoa cthulhu
Pisauridae51356nursery web spidersPisaura mirabilis
Plectreuridae231
Psechridae261
Salticidae6356080jumping spidersZebra spider
Scytodidae5248spitting spidersScytodes thoracica
Segestriidae4130tubeweb spidersSegestria florentina
Selenopidae10257wall spidersSelenops radiatus
Senoculidae131
Sicariidae31623recluse spidersBrown recluse
Sparassidae881224huntsman spidersAvondale spider
Stenochilidae213
Stiphidiidae20125Tartarus mullamullangensis
Symphytognathidae873dwarf orb-weaversPatu digua
Synaphridae313
Synotaxidae111
Telemidae1079long-legged cave spiders
Tetrablemmidae27129armored spiders
Tetragnathidae48996long jawed orb-weaversOrchard spider
Theridiidae1242503cobweb spidersRedback spider
Theridiosomatidae19124ray spidersTheridiosoma gemmosum
Thomisidae1702171crab spidersGoldenrod spider
Titanoecidae553Goeldia obscura
Toxopidae1482
Trachelidae18232
Trechaleidae16120
Trochanteriidae19153
Trogloraptoridae11Trogloraptor marchingtoni
Udubidae415
Uloboridae19283hackled orb-weaversUloborus walckenaerius
Viridasiidae29
Xenoctenidae433
Zodariidae851141ant spidersZodarion germanicum
Zoropsidae26180Zoropsis spinimana

References

Citations

  1. www.publish.csiro.au http://www.publish.csiro.au/samples/spiders/html/SPIDERS.HTM. Retrieved 2022-01-07. {{cite web}}: Missing or empty |title= (help)
  2. Magalhaes, Ivan L. F.; Azevedo, Guilherme H. F.; Michalik, Peter; Ramírez, Martín J. (February 2020). "The fossil record of spiders revisited: implications for calibrating trees and evidence for a major faunal turnover since the Mesozoic". Biological Reviews. 95 (1): 184–217. doi:10.1111/brv.12559. ISSN 1464-7931. PMID 31713947. S2CID 207937170.
  3. Ledford, Joel; Derkarabetian, Shahan; Ribera, Carles; Starrett, James; Bond, Jason E.; Griswold, Charles & Hedin, Marshal (2021). "Phylogenomics and biogeography of leptonetid spiders (Araneae : Leptonetidae)". Invertebrate Systematics. 35: 332–349. doi:10.1071/IS20065.
  4. Coddington, Jonathan A. (2005). "Phylogeny and classification of spiders" (PDF). In Ubick, D.; Paquin, P.; Cushing, P.E. & Roth, V. (eds.). Spiders of North America: an identification manual. American Arachnological Society. pp. 18–24. Retrieved 2015-09-24.
  5. World Spider Catalog (2018), Currently valid spider genera and species.

General sources

Explanatory notes

  1. Unless otherwise shown, currently accepted families and counts based on the World Spider Catalog version 19.0 as of 11 July 2018.[5] In the World Spider Catalog, "species" counts include subspecies. Assignment to sub- and infraorders based on Coddington (2005, p. 20) (when given there).
  2. June 2019 data
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